Good discussion and question. I don’t think the economic counter-incentives will be present in the ‘good’ dApps, since these are usually well thought-out and tested before the network is publicly released. The economic incentives are usually aligned very closely with the token holders, and involve the token holders in the ‘upkeep’ or running of the network.

But to your question, I think like non-dApps, dApps will have many models to choose from. Some that come to mind:

• Sell access to the functionality of their dApp (iExec, Golem, Gnosis, Storj)
• Sell access to their network (BAT, AdEx, Bloom)
• Distribute profits of the product to token holders (TenX, Numerai)

These models are interesting for humans, but they become more interesting when we start thinking of machine to machine transactions, using multiple networks and dApps to achieve their goals.