She was not specific in her article, but the issue seems like one of the candidate choosing to comment on his interviewers’ appearance while being interviewed. That is not the same as a “you look nice today” comment between friends or coworkers or strangers at a bar. This is a job interview. Making statements about your interviewer’s appearance (especially if the candidate did not do anything similar when interviewed by men) is simply unprofessional. Further, we don’t know if the statements were closer to “you look very nice today” or “wow you are smokin’ hot” or “ugh, the last lady was so fat it was disgusting.” No matter what, that was unprofessional and disrespectful. Combine statements concerning the female interviewer’s appearance with the vocalization of doubts about her competence and you have a situation where the interviewer has shown that he will likely not work well with a female boss/colleague. Whatever he said was enough to make the people running the interview uncomfortable, and if they were uncomfortable interacting with him in an interview situation then it is reasonable to extrapolate that they would not feel comfortable working along side of him. This is the purpose of the culture interviews.
And no, it had nothing to do with taking out men. From the sound of it most of the interviews were conducted by men. It seems like there are many men that work there. They were about to hire the guy before he managed to alienate everyone in a single interview. The decision to not hire this particular unnamed man was simply about maintaining a healthy workplace environment.